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According to the most widely accepted hypothesis i.e

Protein assessments by enzyme-linked immunosorbent assay (ELISA) may help to identify mucins directly and having greater utility with mucus samples collected in vivo [74, 84]

Comparison of metallothionein gene expression and nonprotein thiols in tenArabidopsisecotypes (correlation with copper tolerance)

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Mechanistically, increased ECM stiffness activates integrin-mediated mechanotransduction pathways, notably the focal adhesion kinase (FAK), RhoA/ROCK, and YAP/TAZ signaling cascades (151)

doi: 10.1210/me.2003-0048 113 BarthelTKMathernDRWhitfieldGKHausslerCAHopperHATHsiehJCet al